Pulmonary Function and Metabolic Physiology in Scipionyxand other Theropod Dinosaurs

J.A. Ruben, C. Dal Sasso, N.R. Geist, W.J. Hillenius, T.D. Jones, and M. Signore.  1999.
Science 283:514-516.

Abstract: Ultraviolet light analysis reveals that the liver subdivides the visceral cavity of the theropod dinosaur Scipionyx samniticus into distinct anterior pleuro-pericardial and posterior abdominal regions. In addition, Scipionyx apparently possessed diaphragmatic musculature and a dorsally attached posterior colon. These features provide evidence that diaphragm-assisted lung ventilation was present in theropods and that these dinosaurs may have utilized a pattern of exercise physiology distinctly unlike that in any group of living tetrapods.



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